1. In c3 pathway, ribulose biphosphate is the first acceptor of CO2. ADVERTISEMENTS: In this article, we will discuss about the C4 – Dicarboxylic Acid Pathway (Hatch–Slack Pathway). Overview of C4 Pathway Photosynthesis The Calvin cycle was believed to be the only photosynthetic reaction series operating in higher plants and algae for a significant amount of time. (a) The CAM pathway fixes CO{eq}_2 {/eq} onto the three-carbon molecule to make the four-carbon molecule at night, when it is cooler and stomata are open. It is the first step in extracting carbon from carbon dioxide to be able to use it in sugar and other biomolecules. Figure \(\PageIndex{1}\): The C4 Pathway The C4 pathway is designed to efficiently fix CO2 at low concentrations and plants that use this pathway are known as C4 plants. Thus in C4 plants, C4 carbon fixation has a net added cost of 1 ATP for every CO2 delivered to rubisco; however, C4 plants are less likely to die of dehydration compared to C3 plants in dry conditions. They have their stomata open for shorter periods during the day to stop the lose of off too much H2O because of the hot temperatures. It is one of three known processes for carbon fixation “Classic” C\(_4\) plants drive C\(_4\) pathway in leaf mesophyll cells whereas their C\(_3\) is located in so-called bundle sheath cells. 9.7: C4 Pathway and CAM Overview. These plants fix CO2 into a four carbon compound (C4) called oxaloacetate. ). C4 is the alternative pathway of Calvin cycle (C3 cycle) taking place during the dark phase of photosynthesis. 2. C4 carbon fixation or the Hatch-Slack pathway is a photosynthetic process in some plants. But in 1965 Kortschak, Hartt and Burr reported that 4-C containing dicarboxylic acids, […] In c4 pathway, phosphoenol pyruvate is the first acceptor of CO2, while ribulose biphosphate is the second acceptor. Oxaloacetic add is the first product in c4 pathway. For considerable period of time the Calvin cycle as described earlier was thought to be the only photosynthetic reaction sequence operating in higher plants and algae. In the C4 cycle the first stable compound is 4 carbon compound, namely oxaloacetic acid. Hence it is called C4 cycle. It is a process of carbon dioxide fixation. 3. The image below illustrates the pathway we just described: This is a spatial , rather than temporal separation. This occurs in cells called mesophyll cells. Some plants, like sugar cane and corn, that grow in hot conditions, use an alternative process called the C 4 pathway to fix carbon. In c3 pathway, Phosphoglyceric acid is the first product. 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